A row of ToolbarButtons appears to the right of a tiddler title when the mouse is hovered over it. The buttons are:
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close - close the current tiddler
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edit - edit the current tiddler
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fields - show the custom fields attached to the current tiddler
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syncing - shows the server synchronisation status of the current tiddler
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jump - jump straight to another open tiddler
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permalink - puts a link direct to the current tiddler into the address bar
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references - displays all the tiddlers that link to the current tiddler
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done - save changes to a tiddler being editted
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cancel - cancel changes to a tiddler being editted
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delete - delete the current tiddler
The shadow tiddler ToolbarCommands allows these commands to customised and reordered.